Abstract
The beetles of the subfamily Cassidinae s.l. are commonly known as tortoise, presented a world-wide distribution, however they have more diversity in the tropical regions, mainly in South America. Cytogenetically, less than 2% of the Cassidinae species were analyzed, which showed diploid number ranging from 2n=16 to 2n=51, and sex chromosomes system of the Xyp, X0, Xy, Xyc, Xyr, neoXY, Xy…