Abstract
Tool use has been observed in chimpanzees, orangutans, long-tailed macaques and capuchin monkeys, and can present great variation between populations. Capuchins use percussive stone tools to process and access protected resources, such as seeds, fruits and mollusks, and also as "hoes" to dig up tubers, roots and invertebrates from their burrows. To reach for hidden prey, some capuchins' p…